Transforming Monocots Using Agrobacterium

Transforming Monocots Using Agrobacterium Agrobacterium tumefaciens is said to infect dicots naturally. What are the potential obstacles in Agrobacterium-mediated transformation of monocots? Discuss how did the breakthrough (success in transforming monocots using Agrobacterium) come about? (60 marks) Gene transfer using Agrobacterium is a method of transferring genes by using a carrier to insert the gene of interest into the recipient host plant cells. This technology is based on the discovery of infection tumor in the dicotyledone plants caused by a bacterium, named Agrobactertum tumerfaciens. The species Agrobacterium is a soil bacterium which is capable to infect and caused plant wound and then developed into crown galls, normally formed at the trunk of many types of dicot plants. This Agrobactereium spp. has a special DNA, which has a small ring inside the cytoplasm called Ti plasmid (tumour inducing plasmid). On the Ti plasmid, there is a DNA fragment called T-DNA (transfer DNA) which contains the gene causing crown galls development. Plant cells have genes to code for the production of auxin and cytokinin, the two plant hormones which are used as energy sources by Agrobacterium. The use of Ti plasmid in gene transfer into plants is done by replacing the gene related to plan t hormone production and the gene producing opine substance with the desirable trait gene on the T-DNA and then using the Agrobacterium to transfer the gene to the plant chromosomes. Transformation of dicotyledenous plants using Agrobacterium tumefaciens has been well established and widely used but not so in the case of monocotyledonous plants. The potential obstacle in Agrobacterium-mediated transformation of monocot plants includes: Agrobacterium is responsive to phenolic compounds such as acetosyringone which are produced when the plant was wounded. The released phenolic compound from the wounded plant cells will stimulate the performance of vir gene on the Ti plasmid, leading to the transferring T-DNA to the plant chromosome. Most of the dicot plants produced this phenolic compound. On the other hand, most monocot plants did not produce the compounds or produced it in a smaller quantity, therefore resulted in the low efficiency of the Agrobacterium attachment. Furthermore, the wounded cells in the monocot plants multiplied less than in dicot plants. Tissue browning and necrosis following Agrobacterium infection is still a major obstacles especially in cereals. For example in case of wheat, following Agrobacterium infection, wheat embryo and root cells may produce hydrogen peroxide, which altered cell wall decomposition and resulted in a higher level of cellular necrosis and subsequently caused cell death. However the improvement method to resolve the cell death and to improve the transformation efficiency has been demonstrated in cereals (Frame et al., 2002) Apart from necrosis, physical characteristic and genotype, other factors affected transformation efficiency are strains of Agrobacterium used, binary vector, selectable marker gene and promoter, inoculation and co-culture conditions, inoculation and co-culture medium, osmotic treatment, desiccation, Agrobacterium density and surfactants, tissue culture and regeneration medium (Cheng et al., 2004). The Agrobacterium has specificity in attaching monocot plants. Most of monocot plants with important economic value are not hosts of the Agrobacterium, therefore the transformation efficiency involving them is low (Lippincott, 1978). Explants type, quality and source also affect the transformation efficiency foe example embryogenic callus derived from mature seed of rice was reported to be the best explant for Agrobacterium-mediated transformation of rice due to its active cell division (Hiei et al., 1994). The breakthrough on the transformation of monocot plants using Agrobacterium started when Hiei et al. (1994), done a research on Japonica rice. They reported a stable transformation of Japonica rice by using Agrobacterium. They reported results of evaluations using molecular and genetic analysis on the R0, R1 and R2 progenies. The LBA 4404, the super-binary vector of Agrobacterium strain was demonstrated as the most effective vector for the transformation of three Japonica cultivars tested. Their success has open up the possibility of using Agrobacterium for transforming monocot plants such as maize, barley and wheat. In 1996, Ishida et al., has done a transformation research on maize by using a similar approach as developed by Hiei et al (1994). Their transformation efficiency was further improved by the addition of silver nitrate in the culture medium. Other factors that may influence transformation efficiency were also investigated that included incubation time and co-cultivation period. Zhao et al. (2002) optimized the transformation conditions based on Ishidas protocol and it was demonstrated that maize can be transformed with high efficiency by using Agrobacterium method. The gene transfer was done by using a combination of standard binary vector with the addition of antioxidant cysteine in the co-culture medium. In the same year, other researchers included had demonstrated that elite maize cultivars could also be transformed by using Agrobacterium-medated transformation method. Soon after maize, the successful Agrobacterium-mediated transformation of wheat and barley was reported (Jones H.D, 2005, Tingay et al., 1997). Compared with rice and maize, progress with wheat and barley has been slower. Various factors that influence the transformation efficiency have been further investigated. It was reported that the use of surfactant such as Silwett L-77 and desiccation treatment during co-cultivation increased the transformation efficiency of wheat. In the case of barley, since the success of Tingay et al., (1997) in transforming barley by using Agrobacterium, a number of other researchers around the world have reported the successful production of transgenic barley plants. However majority of the successful reports of Agrobacterium-mediated transformation of barley are restricted with model genotype golden promise and igri. Therefore, optimizations of parameters are required to extend the Agrobacterium-mediated transformation in other elite barley cultivars. The transformation of sorghum is the least successfully manipulated. Zhao et al. (2000) developed an efficient Agrobacterium-mediated transformation system for sorghum and from the research it showed that the embryos from the field had higher transformation frequency than those from the greenhouse. Other transformation of monocotyledon plant reported such as Agrobacterium-mediated transformation of turfgrasses, such as creeping bentgrass (Yu et al., 2000), Italian ryegrass (Bettany et al., 2003), and tall fescue (Wang and Ge, 2005) were also reported. Although the delivery of foreign gene into several monocot species via Agrobacterium tumefaciencs has now become a routine technique, there are still serious limitations on the used of this technology on other major monocots. In order to achieve better success in transforming monocot using Agrobacterium, many factors and conditions were being investigated, such as selection of which target tissues which are highly responsive, adjustment of gene transfer conditions to increase the possibility of Agrobacterium attachment into the cell by adding phenolic substances such acetosyringone during co-cultivation period or in co-cultivation medium, that are similar to the substance released by plant cells when they are naturally wounded, using efficient promoter gene to stimulate the expression of the gene in monocot plants and the used of super-virulent of Agrobacterium strains to increase the transformation efficiency (Cheng et al., 2004).

Comparing Twelve Songs by W.H.Auden and Do Not Go Gentle Into That Good

Comparing Twelve Songs by W.H.Auden and Do Not Go Gentle Into That Good Night by Dylan Thomas The two poems I am comparing are 'Twelve Songs' by W.H.Auden and 'Do Not Go Gentle Into That Good Night' by Dylan Thomas. Two themes run through both of these poems, those themes are grief and death. Although the subjects are the same, the poets each have a very different approach to the subjects. This is shown in many different ways, But although their approach to the subjects are very different, their overall views on grief and death are quiet similar. That being that neither grieving nor death is good, and no one wants it to happen. Someone who has lost a loved one, and feels nothing is important now that they have gone is narrating 'Twelve Songs'. A son is narrating 'Do Not Go Gentle Into That Good Night'. The son is telling his father not to die. 'Twelve Songs' is not talking to anyone he is talking about someone. Whereas 'Do Not Go Gentle Into That Good night' the son is talking to his father rather than talking about him. The tone of each poem is quite similar. They both use orders and commands but they use them very differently. 'Twelve Songs' uses a list of instruction or commands throughout the whole poem. 'Do Not Go Gentle Into That Good Night' uses two instructions: "Do Not Go Gentle Into That Good Night" and "Rage rage against the dying of the light". Dylan Thomas uses these two lines throughout the whole poem but only uses those two instructions either at the beginning or the end of each stanza. W.H.Auden uses a lot of Instructions, in fact the whole poem is just instructions:" Stop all the clocks''pack up the moon and dismantle the sun" But none of the instructions are repeated at any other poi... ...ave out that message to the reader. Dylan Thomas did not really give out a message to the reader apart from to put up a fight against death. His aim of begging his father not to die and to create an impact worked. He did use instructions but not in the same way as W.H.Auden, however his instructions did work as he used them to beg his father to 'not go gentle into that good night'. The main difference between these poems is that 'Twelve Songs' is about after death and 'Do Not Go Gentle In to That Good Night' is about before death. The main similarity is that they both use instructions and even though they use them very differently they are still very effective in both poems and are used well. Also both poems talk about personal feelings but only at the end. "I pray do not go gentle into that good night", and "I thought love would last forever I was wrong."

Successes and Failures in Biotechnology Innovation Essay -- Business M

Successes and Failures in Biotechnology Innovation   Ã‚  Ã‚  Ã‚  Ã‚  When I first started this paper I would have to admit that I was pretty green in the field of Biotechnology, I had a brief understanding but nothing near an in depth understanding of the field. So when I first started looking for a success story, I tied my views on successful innovation in the areas that I am familiar with to the biotech field. The major theme that emerged was that successful innovation equals a product that produces quality profits for a company. After further research on the success of innovation, you do realize that profits may be the end goal of the company but success stems from a total company wide effort, that may start many years before a single cent of profit is ever realized.   Ã‚  Ã‚  Ã‚  Ã‚  In my opinion, you have to look at three major areas to see if a company successfully manages their innovation; at the company level, customer level, and the impact the innovation had on society. From the company viewpoint, you have to look at how well management set the company up to be innovators in a given market. Did they choose an industry that would provide the company potential for continuous growth? How did they utilize their employee base and acquire a strong base of knowledge to continually come up with new ideas in the area of interest they are trying to enter? In addition once a product is developed did they manage their intellectual property in the in a manner which provided themselves maximum protection? Finally you have to look at the end results of the innovation. Did they introduce an innovation that allowed the company revenue to grow, were they able to achieve a strong segment of market share, did this innovation have a positive effect on the co mpanies stock price, etc?   Ã‚  Ã‚  Ã‚  Ã‚  From the prospective of the customer you have to look at if the innovation introduced by the company successfully met their needs. Would this product help them save time and money and improve their overall business? How did they manage the channels to which deliver their products to their customers? Your customer will ultimately impact your final sales figures, if they view the product as a benefit to them and if you can meet the needs that the customer has you can have your innovation viewed as a success.   Ã‚  Ã‚  Ã‚  Ã‚  Finally from the standpoint of how the companies innovation had an impact on society. Was... ...sequence can be felt. With the world population soaring you will continue to see the emergence of genetically engineered crops to continue and you will be able to see who the winners and losers are within that industry based on how they handle the introduction of their products. Works Cited Current Status of Starlink Corn. 3 Jun. 2002. Starlinkcorn.com. 2 Feb. 2005.   Ã‚  Ã‚  Ã‚  Ã‚   Estes, Lane. Economic Analysis of Roundup Ready Soybeans. Feb. 2002. 2 Feb. 2005. Geo-Pie Project. Am I eating Soy Beans. 16 Aug. 2004. Genetically Engineered Organisms. 2 Feb. 2005.   Ã‚  Ã‚  Ã‚  Ã‚   Harl, Neil, Roger Ginder, Charles Hurburgh and Steve Moline. The Starlink Situation   Ã‚  Ã‚  Ã‚  Ã‚  25 Oct. 2000. AG Biotech Info Net. 2 Feb. 2005.   Ã‚  Ã‚  Ã‚  Ã‚   Harris, Andrew. GE Corn Pollution Spawns 30 Lawsuits. 18 Sept. 2002. Organicconsumers.org. 2 Feb. 2005. Monsanto Inc. Simply the Most Profitable Way to grow Soy Beans. Jan. 2005. Monsanto Inc. 2 Feb. 2005. Ruen, Jim. Biotech R&D Isn’t Just About Traits. Jan. 2005. Dealer and Applicator. 2 Feb. 2005. Starlink Corn. FSANZ. 2 Dec. 2004. Foodstandards.gov. 2 Feb, 2005. US 2000/01 Corn Exports Dented by Starlink. 10 Sept. 2001. Planetark.com. 2 Feb 2005.

Comparing and Contrasting the Children in Susan Skinner’s ‘the Minnow Catching Boys’ and ‘My Parents Kept Me from Cildren Who Were Rough’ by Stephen Spender

Comparing and Contrasting the children in Susan Skinner’s ‘The Minnow Catching Boys’ and ‘My Parents Kept Me from Children who were Rough’ by Stephen Spender. In this essay I am going to discuss ‘The Minnow Catching Boys’ by Susan Skinner and ‘My Parents kept me from Children who were Rough’ by Stephen Spender. I will focus on the similarities and differences between the children in the poems in terms of their actions, the language used to describe them and their impact on the reader. The Minnow Catching Boys’ is a poem about a group of boys who very skillful and adventurous. ‘My Parents kept me from children who were rough’ is also about a group of boy bullies who really torture others a lot and play rough. The children in ‘The Minnow Catching Boys’ are described as skillful and the outdoor-type boys. This poem is based on a Victorian/older lifetime according to its facts; Boys nowadays play video games and are interested in technology or sports. This also refers to the quote â€Å"the sun burnt boys’ which means that sunscreen was not invented.I would imagine them as scruffy boys wearing a shirt with a waistcoat and knee breeches (knee length shorts from older times); all dull colours and by looking back it says ‘Breeches rolled’. The children are in a focused, competitive mood, as said in the poem, ‘Scan and dip’ as well as ‘they make bets poke and tip, their nimble catch’. ‘For they have always been here the minnow catching boys. ’ makes me think that this competitive hobby has been passed down from many generations. ‘†¦Ã¢â‚¬ ¦.. And when they have had enough, the minnow catching boys make their way home’ makes me think these boys are very carefree.I thought quotes like ‘†¦Ã¢â‚¬ ¦Ã¢â‚¬ ¦. And a small girl who can never quite catch up calls for them to wait, but no one waitsâ€⠄¢ give me a feeling that these boys are mean and cruel or maybe just in a rush. The quote ‘ The dog shakes off in a cascade of diamonds’ is one of the most striking pieces of imagery for me because it imaginatively describe thousands of sparkling water droplets as a ‘cascade of diamonds’. The tone which this poem would be read in would be slow and suspending- long pauses. You would read this poem in this tone so you can get your audience right ‘into’ the scene-as the boys are focused and concentrating too hard.The children in ‘My Parents kept me from Children who were rough’ are described as mean, arrogant and a fierce group of bullies. I imagine them as very scruffy boys with their clothes torn; as they don’t care. I think they would be wearing the same clothes as the boys in the other poem but in an ill-suited manner. I think this poem is also set in the same time as the other poem- older times/Victorian times. This poem quotes ‘They threw words like stones’ which makes me think that these boys are rude, ill-mannered and that their insults really ‘hurt’ people- using words like weapons.These boys were also involved in physical abuses to others, which refers to the quote’ Their knees tight on my arms’. ‘They were lithe’, this quote makes me think these boys are super-fit, strong, flexible and very versatile. I think these boys look like are big headed and want to be threatening to people. Their parents might not able to control them or maybe are not even aware of their actions. ‘Muscles like iron ‘says that these boys are unbeatable and formidably strong. I would like to read this poem in a fast, envious tone, because I think that the narrator is envying these strong and super-fit boys. Comparing and Contrasting the Children in Susan Skinner’s ‘the Minnow Catching Boys’ and ‘My Parents Kept Me from Cildren Who Were Rough’ by Stephen Spender Comparing and Contrasting the children in Susan Skinner’s ‘The Minnow Catching Boys’ and ‘My Parents Kept Me from Children who were Rough’ by Stephen Spender. In this essay I am going to discuss ‘The Minnow Catching Boys’ by Susan Skinner and ‘My Parents kept me from Children who were Rough’ by Stephen Spender. I will focus on the similarities and differences between the children in the poems in terms of their actions, the language used to describe them and their impact on the reader. The Minnow Catching Boys’ is a poem about a group of boys who very skillful and adventurous. ‘My Parents kept me from children who were rough’ is also about a group of boy bullies who really torture others a lot and play rough. The children in ‘The Minnow Catching Boys’ are described as skillful and the outdoor-type boys. This poem is based on a Victorian/older lifetime according to its facts; Boys nowadays play video games and are interested in technology or sports. This also refers to the quote â€Å"the sun burnt boys’ which means that sunscreen was not invented.I would imagine them as scruffy boys wearing a shirt with a waistcoat and knee breeches (knee length shorts from older times); all dull colours and by looking back it says ‘Breeches rolled’. The children are in a focused, competitive mood, as said in the poem, ‘Scan and dip’ as well as ‘they make bets poke and tip, their nimble catch’. ‘For they have always been here the minnow catching boys. ’ makes me think that this competitive hobby has been passed down from many generations. ‘†¦Ã¢â‚¬ ¦.. And when they have had enough, the minnow catching boys make their way home’ makes me think these boys are very carefree.I thought quotes like ‘†¦Ã¢â‚¬ ¦Ã¢â‚¬ ¦. And a small girl who can never quite catch up calls for them to wait, but no one waitsâ€⠄¢ give me a feeling that these boys are mean and cruel or maybe just in a rush. The quote ‘ The dog shakes off in a cascade of diamonds’ is one of the most striking pieces of imagery for me because it imaginatively describe thousands of sparkling water droplets as a ‘cascade of diamonds’. The tone which this poem would be read in would be slow and suspending- long pauses. You would read this poem in this tone so you can get your audience right ‘into’ the scene-as the boys are focused and concentrating too hard.The children in ‘My Parents kept me from Children who were rough’ are described as mean, arrogant and a fierce group of bullies. I imagine them as very scruffy boys with their clothes torn; as they don’t care. I think they would be wearing the same clothes as the boys in the other poem but in an ill-suited manner. I think this poem is also set in the same time as the other poem- older times/Victorian times. This poem quotes ‘They threw words like stones’ which makes me think that these boys are rude, ill-mannered and that their insults really ‘hurt’ people- using words like weapons.These boys were also involved in physical abuses to others, which refers to the quote’ Their knees tight on my arms’. ‘They were lithe’, this quote makes me think these boys are super-fit, strong, flexible and very versatile. I think these boys look like are big headed and want to be threatening to people. Their parents might not able to control them or maybe are not even aware of their actions. ‘Muscles like iron ‘says that these boys are unbeatable and formidably strong. I would like to read this poem in a fast, envious tone, because I think that the narrator is envying these strong and super-fit boys.

Electronic medical records Essay

Electronic medical records are believed to be the way of the future. Hospitals and other healthcare settings are increasingly turning to electronic records over traditional paper records. However, many still have not made the leap and continue to use paper instead of electronic. Healthcare practices must weigh the pros and cons before deciding which records management system to use. An electronic health record (EHR) is a representation of all a patients’ data; know as a digital version of a patient’s paper chart. Paper-based records are the most common method of recording patient information for most doctor’s offices and hospitals in the United States. The digital information is usually stored in a database and is accessible from everywhere via a network and EMRs contain mainstream data normally found on a patient’s medical records. It contains all information ranging anywhere from a patient’s medical history, diagnoses, medications, treatment plans, immunization dates, allergies, radiology images, and laboratory and test results that have been combined and structured in a digital form. It allows for an entire patient history to be viewed without the need to track down the patient’s previous medical record volume and assist in ensuring data is accurate, appropriate and legible. It reduces the chances of data replication, as there is only one modifiable file, which means the file is constantly up to date when viewed at a later date and eliminates the issue of lost forms or paperwork. There have been many issues debating if this is a good system, and pros and cons that go hand in hand. Electronic records have many benefits, including accessibility. They are currently the preferred system because of how easily they make it for doctors to coordinate patient care. Accessing electronic records is a lot easier and faster than waiting to receive paper ones. This can greatly speed up doctor collaborations in patient care and perhaps improve the quality of care that patients receive. There are many companies that sell these systems, so healthcare practices can choose a system with an EMR interface and features that best suit their needs. These interfaces can also be coordinated with billing systems for an extra degree of convenience and functionality. The case for medical records is compelling. They can make healthcare more efficient and less expensive. It can also improve the quality of our healthcare system by making patients medical history more easily accessible to all the people who treat them, being a hospital or doctor. The Government has given 6.5 million in incentives and hospitals and doctors have spent billions more. Some complain that the electronic systems are time consuming and designed more for bureaucrats than physicians. Although the health care industry has been slower than many others to replace paper records with electronic ones, some of the advantages of computerized systems are becoming obvious. EMRs translate into better treatment for patient’s efficiency and speed of diagnosis translates into better health care service for patients. Similar to the previously discussed point, correct and timely information can significantly increase the quality of health care service rendered to patients. EMRs can save lives. Improvements to patient safety are found throughout EHRs. For one, drug-drug and drug-allergy checks help prevent unintended reactions to medications. Secondly, photo-enabled EHRs help identify the patient. Thirdly, but definitely not the last one, barcode medication administration checks for the five rights to ensure the right patient receives the right medication at the right time. Electronic health records provide an avenue for better care coordination between multiple health care providers and the patient. Information can easily be shared between the inpatient, outpatient and ED settings. Furthermore, patients are able to more easily access their information, either by receiving an electronic summary of care record or logging into a patient portal. Electronic health records provide the tools to help physicians and nurses make better care decisions. For example, clinical decision support tools, like making medication recommendations based on diagnosis, and built-in clinical pathways, which help guide a hospital through recommended procedures, help ensure patients receive the best, research-proven care. EMRs give patients control over their health records. Based on the principle that since it’s the patient’s medical record, the patient should control it, decide what should be in it, and who gets access to it there are currently many online EMRs available. In spite of the many perks, electronic health records do come with some drawbacks. One major disadvantage is the significant start-up costs. From the technology, including hardware installations and software upgrades, to staff training, the start-up costs of migrating to electronic medical records are significant, especially for a small medical practice or for a large medical network of remote offices. There is a lot of time and labor involved up-front to migrate paper files to a completely electronic platform. Another would be the lack of quality patient time. When doctors, nurses, and administrative staff are unfamiliar with the technology and how a new system works, they often spend more time on it. Or, they may be uncomfortable using it so it will take them longer to execute a task. All this lost time could be potentially spent servicing patients or tending to other mission critical matters. Since electronic medical records, as an industry, is still in its infancy, we have yet to see a standardization of EMR requirements and utilization across healthcare organizations, insurance companies, pharmacies, etc. The problem is when these disparate systems do not synchronize; it results in errors, duplication of efforts, or a lag in time in service. Security concerns are still an issue. While many might think that migrating to an electronic system would be safer, another one of the disadvantages of electronic medical records is vulnerability those systems inherently create. There is still a real concern that no matter how many firewalls, password encryptions, and other security features added, there will be a hacker who can beat the system and potentially compromise and steal personal information. All these concerns can be addressed by potentially partnering with an outsource company that specializes in electronic medical records as they have the technology and trained staff to handle large volumes of patient information. They have also invested the time and resources into the latest equipment and security measures. The irony is that while EMRs were designed to improve communication and increase time spent with patients, they actually have enabled some doctors to engage less with patients because they have much more information before actually seeing the patient. In addressing these concerns, there is one key way to circumvent these and other disadvantages of electronic medical records. Working with a highly specialized electronic medical records solutions provider will assure that best-of-breed technologies, and the industry and operational expertise, is in place. By outsourcing the entire process, your staffs can also then focus on core business areas and foster the company’s service reputation. An electronic medical records partner also best ensures systems integrate with others in synergistic industries, and that the best and most current security mechanisms are deployed to keep patient information inaccessible to those unauthorized to view.

High-Growth Industry Jobs Worth Going Back to School

Computer And Video Games Friends Needed - 974 Words

Computer and Video Games: Friends Needed Video games have existed for much longer than many young people are aware. For example, an individual would need to be over the age of 50 to remember the first time arcade games were available for play in a bowling alley or inside the entertainment building of a miniature golf course. Similarly, an age requirement of 40 would be required for a person to remember the first release of the Atari, which was usually seen at a friend’s home, or to remember the first complex games available on a computer (for example, The Sims). With a history of decades, computer and video games have influenced generations. Although many experts and doctors have argued otherwise, computer and video games usage doesn’t encourage or cause violent behavior; rather, participation in computer/video games encourages positive social interaction and strengthens problem-solving skills. Computer games have taken the blame for many national tragedies, including C olumbine and Sandy Hook; however, in the 1970s, rock music was blamed for many violent acts by teenagers, including suicide. Instead of delving into their child’s psychological status or the quality of his/her home life, parents look around for any other reason for their child’s violent behavior that won’t require them to take the blame for the child’s actions. Even the parents of children who weren’t involved exhibit his behavior; the reason could be that it gives comfort for people to think, â€Å"My childShow MoreRelatedVideo Games : A Huge Part Of Our Society900 Words   |  4 PagesVideo games are a huge part of our society more now than ever before. A huge number of people around the world play them. In fact, the game StarCraft boasts to be South Koreas unofficial national sport due to the high number of players. Throughout media and just day-to-day life. 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